Genshin Impact

Dendroculus Locations and Interactive Map

Last updated on: March 13, 2026 03:49 AM

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Genshin Impact - Dendroculus Locations and Interactive Map

There are 271 Dendroculus in Genshin Impact. Here's an interactive map of all Dendroculus locations in Sumeru's Rainforest, Desert, Hadramaveth, and Girdle of the Sands areas!

Elemental Oculus Items
All Oculi	Anemoculus	Geoculus	Electroculus
Dendroculus	Hydroculus	Pyroculus	Lunoculus

List of Contents

Interactive Map and Locations
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Dendroculus Interactive Map and Locations

Dendroculus Locations Map

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Item	Area	How to Get
Dendroculus 1	Sumeru Rainforest	Location: Available in The Final Chapter Quest of Agnihotra Sutra quest chain. Go to the final cave and head to the farthest location and you should see the rock below. Hit the rock with any Dendro attacks. The rock will rise, and the Dendroculus will become accessible.
Dendroculus 2	Sumeru Rainforest	Location: Located in Matiyama Forest inside a giant mushroom/tree.
Dendroculus 3	Sumeru Rainforest	Location: Located northernmost part of Sumeru, on top of a giant mushroom. Use Dendro or get the Dendrogranum and shoot at the three floating targets to remove the barrier to get the Dendroculus.
Dendroculus 4	Sumeru Rainforest	Location: In Mawtiyima Forest, activate the mushroom on the ground close to some Anemo Slimes to summon a wind current and get the Dendroculus high up in the sky.
Dendroculus 5	Sumeru Rainforest	Location: Located on a mushroom inside a cave beneath the waypoint. The tunnel entrance is found southwest of the waypoint.
Dendroculus 6	Sumeru Rainforest	Location: Located in Matiyama Forest on top of a giant mushroom/tree.
Dendroculus 7	Sumeru Rainforest	Location: Located south of Mawtiyima Forest, on top of a giant boulder.
Dendroculus 8	Sumeru Rainforest	Location: Located at the entrance of the cave that leads to Electro Regisvine.
Dendroculus 9	Sumeru Rainforest	Location: Located on top of an umbrella on the hill where the Electro Regisvine's cave is located. Use Dendro on the nearby Clusterleaves of Cultivation to summon a Four-Leaf sigil and get the Dendroculus.
Dendroculus 10	Sumeru Rainforest	Location: Located north of Gandha Hill, between some overhanging roots. Use the Four-Leaf sigil to get it.
Dendroculus 11	Sumeru Rainforest	Location: Located at Lokapala Jungle, south of Mawtiyima Forest on top of a large mushroom with three Anemo Slimes. Use Four-Leaf sigils to get to the top of the mushrooms and jump from one to another.
Dendroculus 12	Sumeru Rainforest	Location: Located at the top of the Palace of Alcazarzaray, Climb the palace or you can go from the northeast teleport waypoint and glide from the cliffs.
Dendroculus 13	Sumeru Rainforest	Location: By the waterfalls west of Gandha Hill, jump from the top of the waterfalls or use any of the Bouncy Mushrooms around the waterfalls to get the Dendroculus.
Dendroculus 14	Sumeru Rainforest	Location: Located under a ledge. Jump and glide down to get the Dendroculus.
Dendroculus 15	Sumeru Rainforest	Location: Located at Chatrakam Cave. Only accessible during Risen Moon Chapter quest under Agnihotra Sutra quest chain. Requires upgraded Kusava gadget.
Dendroculus 16	Sumeru Rainforest	Location: Located inside the cave during Agnihotra Sutra. Hit all floating rings with Dendrograna to break the "barrier."
Dendroculus 17	Sumeru Rainforest	Location: Can only be accessed from the Agnihotra Sutra, Risen Moon Chapter quest.
Dendroculus 18	Sumeru Rainforest	Location: Apply Electro on the Bouncy Mushroom to make Four-Leaf Sigils appear. Jump on mushroom and use the sigils to reach the Dendroculus.
Dendroculus 19	Sumeru Rainforest	Location: Clear the Withering first, then apply Electro on the Bouncy Mushroom to reach the Dendroculus.
Dendroculus 20	Sumeru Rainforest	Location: Located on top of a tree. Use the Four-Leaf Sigils to reach it quickly.
Dendroculus 21	Sumeru Rainforest	Location: Located inside a cave southeast of Vanarana. You'll need to unflood the area by destroying two Withering Branches, then activating the Dendro Monument to reduce the water level.
Dendroculus 22	Sumeru Rainforest	Location: Located in an underground cave, west of Sumeru City past the nearest waypoint and a Withering Zone. Find the tunnel east of the waypoint and follow the path until you find the Dendroculus.
Dendroculus 23	Sumeru Rainforest	Location: Located in a cave full of Hydro Slimes south of Vanarana, near a Statue of The Seven.
Dendroculus 24	Sumeru Rainforest	Location: Located by the road south of Vanarana inside a tent.
Dendroculus 25	Sumeru Rainforest	Location: Located southeast of a Statue of Seven in Vanarana in a little clearing above a pond. Use the Four-Leaf Sigil to reach it, or glide down if you can get to on a higher area.
Dendroculus 26	Sumeru Rainforest	Location: Located just before the waterfall in Vanarana in a little hut.
Dendroculus 27	Sumeru Rainforest	Location: Located inside the hollow space of the big trunk. Use the Four-Leaf Sigils so you can easily get up this area
Dendroculus 28	Sumeru Rainforest	Location: Located in a cave where the little farm is. Enter the cave then go right on the fork.
Dendroculus 29	Sumeru Rainforest	Location: Located by the Jadeplume Terrorshroom boss location, underneath a stone arc that leads to a group of Shroom-Kin enemies.
Dendroculus 30	Sumeru Rainforest	Location: Located south of the Jadeplume Terrorshroom boss location (located at the very northwest of the map). It's floating above the hole in the ground that leads down to the Jadeplume.
Dendroculus 31	Sumeru Rainforest	Location: Located under a rock. Get rid of The Withering zone to unearth it.
Dendroculus 32	Sumeru Rainforest	Location: Located above a waterfall. Use the Bouncy Mushroom on one side of the waterfall to reach the Dendroculus.
Dendroculus 33	Sumeru Rainforest	Location: Located north of Yasna Monument, high up in the sky. Use Electro on the Bouncy Mushroom on top of the large boulder with a Hilichurl Camp built around it to jump high enough to get the Dendroculus.
Dendroculus 34	Sumeru Rainforest	Location: Get rid of the Withering zone to revitalize the Bouncy Mushroom. Use Electro to reach higher!
Dendroculus 35	Sumeru Rainforest	Location: Located inside the cave where you need to lower the water level.
Dendroculus 36	Sumeru Rainforest	Location: Located underground, during the Aranayaka Part III quest.
Dendroculus 37	Sumeru Rainforest	Location: Located north of Pardis Dhyai underneath a rock. It will appear after clearing the Withering zone.
Dendroculus 38	Sumeru Rainforest	Location: Solve all the puzzles inside the cave until you get to the flooded underground area. On the other side, you'll see the Dendroculus floating on top of the trunk. You'll need songs from the Vintage Lyre.
Dendroculus 39	Sumeru Rainforest	Location: Located south of Yazadaha Pool, underneath a stone bridge with large roots. Jump from the top of the bridge to get the Dendroculus.
Dendroculus 40	Sumeru Rainforest	Location: Located left from Yazadaha Pool, high up in the air. Use the Bouncy Mushroom directly under to reach it; to jump higher, use Electro.
Dendroculus 41	Sumeru Rainforest	Location: Located near Yazadaha Pool, high up in the sky, Get rid of The Withering zone on the ground to revitalize the Bouncy Mushroom beneath it. Use Electro on the mushroom to gain more height.
Dendroculus 42	Sumeru Rainforest	Location: Located west of Sumeru city, at the end of the big tree roots. Use the Bouncy Mushroom directly below it; use Electro to jump higher.
Dendroculus 43	Sumeru Rainforest	Location: Located at the tip of one of the branches of the giant tree in Sumeru City. Climb up the side of the Sanctuary of Surastana and onto the tree trunk to get up to the branch.
Dendroculus 44	Sumeru Rainforest	Location: Located under the large wood bridge southwest of Chinvat Ravine; jump from the bridge itself or use the Bouncy Mushroom found at the side to gain height and glide towards it.
Dendroculus 45	Sumeru Rainforest	Location: Located past the large wood bridge underneath a dendro barrel; explode the Dendro barrel with Pyro to unearth it.
Dendroculus 46	Sumeru Rainforest	Location: Located west of Gandharva Ville inside a tree trunk house.
Dendroculus 47	Sumeru Rainforest	Location: Located on top of the giant tree house at Gandharva Ville. Follow the path up and then climb the tree trunk.
Dendroculus 48	Sumeru Rainforest	Location: Located on top of a rocky arch by the road, at the northeast entrance of Gandharva Ville.; can be obtained with Venti/Kazuha or using Geo Constructs.
Dendroculus 49	Sumeru Rainforest	Location: Complete the Agnihotra Sutra quest to get the "Kusava" gadget. Fix the broken monument to clear the roots.
Dendroculus 50	Sumeru Rainforest	Location: Located on top of an entangled boulder.
Dendroculus 51	Sumeru Rainforest	Location: Located under a rock after clearing the Withering.
Dendroculus 52	Sumeru Rainforest	Location: Located inside a cave that is unlocked during the Vimana Agama: Rojinyan's Chapter quest.
Dendroculus 53	Sumeru Rainforest	Location: Located at the edge of a cliff south of Vimara Village, with a Hilichurl Shooter standing right below it.
Dendroculus 54	Sumeru Rainforest	Location: Located inside a ruined hut at the top of the hill.
Dendroculus 55	Sumeru Rainforest	Location: Located on a large gear under a rock arch.
Dendroculus 56	Sumeru Rainforest	Location: Located underwater in an underground cave. Grab it without draining the water by doing a plunging attack from the branch above.
Dendroculus 57	Sumeru Rainforest	Location: Located close to the border of Liyue and Sumeru, southwest of the teleport waypoint by Tiangong Gorge.
Dendroculus 58	Sumeru Rainforest	Location: Summon Dendrograna from the nearby Auspicious Branches and shoot the floating target with a Charged Bow Attack. Do the same to the three targets it summons to lower the barrier.
Dendroculus 59	Sumeru Rainforest	Location: Located on a mound beside a tree.
Dendroculus 60	Sumeru Rainforest	Location: Play the Rhythm of the Beastly Trail using the Vintage Lyre on the flower to reveal the Dendroculus. You can also get it by touching or standing on the rock.
Dendroculus 61	Sumeru Rainforest	Location: Located on a tree east of Devantaka Mountain.
Dendroculus 62	Sumeru Rainforest	Location: Located on top of a gear in the middle of Devantaka Mountain. Glide towards it from the nearest Statue of the Seven (Ardravi Valley, Sumeru).
Dendroculus 63	Sumeru Rainforest	Location: Located inside one of the fingers of the massive corroded robot.
Dendroculus 64	Sumeru Rainforest	Location: Located east of Devantaka Mountains inside a hollowed tree trunk.
Dendroculus 65	Sumeru Rainforest	Location: Located south of Devantaka Mountain under the arm of the massive corroded robot.
Dendroculus 66	Sumeru Rainforest	Location: Located inside the giant robot's eye. Run along its arm to reach the Dendroculus or glide down from the waypoint on the south.
Dendroculus 67	Sumeru Rainforest	Location: Located at the southeastern mountain range of Sumeru, east of Devantaka Mountains inside a Dendro dome. Pick up the flowers, which will spawn Whopperflowers around the Dendroculus and lower the barrier.
Dendroculus 68	Sumeru Rainforest	Location: Located at the southeast area of Devantaka Mountain. It is hidden in tree leaves, and guarded by an enemy. Scale the mountain and get to the top.
Dendroculus 69	Sumeru Rainforest	Location: Located in an underground area south of Devantaka Mountain. Enter the area from the north and head all the way south until you find giant metal parts. Apply Electro on the Bouncy Mushroom and use it to reach the Dendroculus tucked atop of one of the metal objects.
Dendroculus 70	Sumeru Rainforest	Location: Located east of Port Ormos inside a Dendro dome. Once you've unlocked the Rhythm of the Beastly Trail, play it to unlock the dome.
Dendroculus 71	Sumeru Rainforest	Location: Located at the mouth of a river north of Port Ormos. It is on a net beside an inactive Ruin Guard
Dendroculus 72	Sumeru Rainforest	Location: Located in the middle of the lake north of Port Ormos. Grab a waverider from the nearby waypoint to reach it easily.
Dendroculus 73	Sumeru Rainforest	Location: Located on a large gear north of Port Ormos.
Dendroculus 74	Sumeru Rainforest	Location: Located inside a hollow trunk west of Port Ormos.
Dendroculus 75	Sumeru Rainforest	Location: Located on the roof of a hut northwest of Port Ormos.
Dendroculus 76	Sumeru Rainforest	Location: Located inside a giant log southeast of Apam Woods.
Dendroculus 77	Sumeru Rainforest	Location: Clear the Withering southwest of Vimara Village and find the bouncy mushroom right below the Dendroculus. energize it with Electro and the bounce should shoot you straight to it
Dendroculus 78	Sumeru Rainforest	Location: Located in an alcove at the base of the valley in between Pardis Dhyai and Vimara Village, with a Hilichurl digging on top of it.
Dendroculus 79	Sumeru Rainforest	Location: Located beneath a large hollow tree trunk overlooking the river between Pardis Dhyai and Vimara Village. Climb the tree trunk to get the Dendroculus.
Dendroculus 80	Sumeru Rainforest	Location: Located under the bridge south of the Pardis Dhyai. Glide down to get it.
Dendroculus 81	Sumeru Rainforest	Location: Located under a rock after clearing the Withering zone along the path north of Apam Woods.
Dendroculus 82	Sumeru Rainforest	Location: Located at the base of a tree at the entrance of a Withering zone in Apam Woods.
Dendroculus 83	Sumeru Rainforest	Location: Located in the middle of a waterfall in Apam Woods.
Dendroculus 84	Sumeru Rainforest	Location: Located under a rock after clearing a Withering zone east of Apam Woods.
Dendroculus 85	Sumeru Rainforest	Location: Beneath a tree at the center of Apam Woods. Go up the slope on the west side and glide to reach it.
Dendroculus 86	Sumeru Rainforest	Location: At the top of a waterfall in Apam Woods, use the nearby Clusterleaves of Cultivation to release a Four-Leaf Sigil that can help you reach this Dendroculus.
Dendroculus 87	Sumeru Rainforest	Location: From the Tower of Abject Pride domain, glide east and land on a platform on the side of the tree. Use the Bouncy Mushroom and the Four-Leaf Sigils to reach a higher platform. Use Dendro on the Clusterleaves of Cultivation to create a trail of sigils that will lead to this Dendroculus.
Dendroculus 88	Sumeru Rainforest	Location: Located under a plant after clearing the Withering zone north of the Tower of Abject Pride domain.
Dendroculus 89	Sumeru Rainforest	Location: Found after completing the four subquests of the Varuna Gatha quest chain. Located south of Apam Woods on an island in the middle of the lake.
Dendroculus 90	Sumeru Rainforest	Location: Located on a large tree southeast of Apam Woods. Use the Bouncy Mushroom near the teleport waypoint for a boost up.
Dendroculus 91	Sumeru Rainforest	Location: Located under a giant root north of The Coordinates of Sun and Rain domain. Climb the root and glide down to reach the Dendroculus.
Dendroculus 92	Sumeru Rainforest	Location: Located under a plant after clearing the Withering zone southwest of The Coordinates Sun and Rain domain.
Dendroculus 93	Sumeru Rainforest	Location: Located under a wooden platform southeast of the Tower of Abject Pride domain.
Dendroculus 94	Sumeru Rainforest	Location: Located in Grove of Dreams (underground area), on the topmost branch of the large tree.
Dendroculus 95	Sumeru Rainforest	Location: Located east of Ruins of Dahri, under a plant holding a rock. Clear the Withering zone first to reveal the Dendroculus.
Dendroculus 96	Sumeru Rainforest	Location: Located down the hole in Ruins of Dahri. You can only access this by playing the Irate Iron Chunk quest of Varuna Gatha.
Dendroculus 97	Sumeru Rainforest	Location: Located south of Ruins of Dahri, near the teleport waypoint. Carefully climb the overgrowth to the tip of one of the roots.
Dendroculus 98	Sumeru Rainforest	Location: Play Rhythm of the Gloomy Path (learned from The Rhythm that Leads to the Gloomy Path quest). Enter the portal to be transported to the Dendroculus.
Dendroculus 99	Sumeru Rainforest	Location: Located at the top of the giant tree in Ruins of Dahri.
Dendroculus 100	Sumeru Rainforest	Location: Play Rhythm of the Gloomy Path (learned from The Rhythm that Leads to the Gloomy Path quest). Enter the portal. You'll be transported to the Dendroculus.
Dendroculus 101	Sumeru Rainforest	Location: Located under a rock after clearing a Withering zone north of Caravan Ribat.
Dendroculus 102	Sumeru Rainforest	Location: To get this, do the quest The Rhythm that Nurtures the Sprout. Unlock the Dendrograna branch and get a Healthy Dendrograna by using the Vintage Lyre and shoot the floating target.
Dendroculus 103	Sumeru Rainforest	Location: Located on top of a tower to the left of Yasna Monument. Glide down from the closest Teleport Waypoint to reach it.
Dendroculus 104	Sumeru Rainforest	Location: Located in Yasna Monument. Revive the Dendrograna Branch with the Rhythm of the Sprout, then hit all floating Dendro rings behind the rock piles beyond the big branches with the Healthy Dendrograna.
Dendroculus 105	Sumeru Rainforest	Location: Located above a stone statue on top of a hill.
Dendroculus 106	Sumeru Rainforest	Location: Get rid of the Withering zone to unearth the dendroculus from beneath the stone
Dendroculus 107	Sumeru Rainforest	Location: Get rid of the Withering zone to unearth the Dendroculus from beneath the stone
Dendroculus 108	Sumeru Rainforest	Location: Located north of Yasna Monument on top of a large tree.
Dendroculus 109	Sumeru Rainforest	Location: Complete For All Children Who Long for Life so you can safely explore Old Vanarana from the effects of the Withering. Apply Dendro to his rock, then the rock will spring up to reveal the Dendroculus.
Dendroculus 110	Sumeru Rainforest	Location: Complete For All Children Who Long for Life so you can safely explore Old Vanarana from the effects of the Withering. The Dendroculus is on top of the spiral dead tree.
Dendroculus 111	Great Red Sand	Location: Located west of the Eye of the Sand. It is above the gate next to a teleport waypoint.
Dendroculus 112	Great Red Sand	Location: Located right above a remarkable chest. The area can only be accessed during the Dreams Beneath the Searing Sand World Quest.
Dendroculus 113	Great Red Sand	Location: Located in the same room as an Obscuring Cell and a Primal Beam during the Dreams Beneath the Searing Sand World Quest. The Dendroculus is found behind the crumbled wall on the north side of the room.
Dendroculus 114	Great Red Sand	Location: Located at the end of the invisible maze inside Khaj-Nisut.Only accessible during the Dreams Beneath the Searing Sand World Quest.
Dendroculus 115	Great Red Sand	Location: Only accessible after revealing Khaj-Nisut during the Dreams Beneath the Searing Sand World Quest. Glide down from the southeast waypoint to the top of Khaj-Nisut to reach this Dendroculus.
Dendroculus 116	Great Red Sand	Location: Located behind a statue in the invisible maze room. The area can only be accessed during the Dreams Beneath the Searing Sand World Quest.
Dendroculus 117	Great Red Sand	Location: Located east of the Eye of the Sand, at the top of the rock formation.
Dendroculus 118	Great Red Sand	Location: Located east of The Dune of Carouses. It is behind the three wooden boxes below the cliff.
Dendroculus 119	Great Red Sand	Location: Located by Abdju Pit, west of a small camp. found by following the Golden Slumber World Quest.
Dendroculus 120	Great Red Sand	Location: Located near a common chest circled by Geo Fungi and a Geoshroom.
Dendroculus 121	Great Red Sand	Location: At the hole that leads underground, drop down the gale to grab it.
Dendroculus 122	Great Red Sand	Location: Located in front of a ruined tower in Aaru Village.
Dendroculus 123	Great Red Sand	Location: Located at the top of the building at the center of Aaru Village, next to the teleport waypoint
Dendroculus 124	Great Red Sand	Location: Located by a hole in Dar al-Shifa.
Dendroculus 125	Great Red Sand	Location: After gaining enough clearance through the Golden Slumber Questline, return to the ruins that you found near Abdju Pit and activate the plinth inside. Follow the path until you find the room with the Primal Construct: Reshaper. The Dendroculus is along a path past this room.
Dendroculus 126	Great Red Sand	Location: Located in the ruins during the Lost in the Sands quest.
Dendroculus 127	Great Red Sand	Location: Found deeper in the ruins near Abdju Pit.
Dendroculus 128	Great Red Sand	Location: Inside the Plint of the Secret Rites room during the Lost in the Sands quest. The entrance is located from the hidden waypoint near the Domain on the top left.
Dendroculus 129	Great Red Sand	Location: Keep following the path that you took while doing the Lost in the Sands quest until you find a second plinth. Operate this plinth after gaining enough clearance to call down an elevator that will help you reach this Dendroculus.
Dendroculus 130	Great Red Sand	Location: Located at the top of an Ajilenakh Nut tree. Use the Four-Leaf Sigil nearby for a boost and glide down to the Dendroculus.
Dendroculus 131	Great Red Sand	Location: Destroy the rocks beneath Dendroculus #14 to find an underground cave. Follow the path, breaking more rock piles along the way, until you find a small Hilichurl camp. Destroy the rock pile behind the camp to find a room containing the Dendroculus.
Dendroculus 132	Great Red Sand	Location: Located just below a cliff east of the Garden of Endless Pillars domain.
Dendroculus 133	Great Red Sand	Location: Jump down the pit next to the waypoint south and find an opening beneath the statue. Follow the path north until you get inside the tower. At the very top, the Dendroculus can be found behind the pedestal.
Dendroculus 134	Great Red Sand	Location: Go inside The Dune of Carouses and change the Prism of Khaj-Nisut's color to blue. This will disable the invisible barrier of the room to your left, where the Dendroculus is.
Dendroculus 135	Great Red Sand	Location: Ride the upward draft south of The Dune of Carouses and follow the trail of wind hoops it creates to reach this Dendroculus.
Dendroculus 136	Great Red Sand	Location: Located southeast of The Dune of Carouses below a ledge northwest of a Statue of the Seven.
Dendroculus 137	Great Red Sand	Location: Glide on the upward draft near the teleport waypoint to reach the Dendroculus just above an invisible barrier.
Dendroculus 138	Great Red Sand	Location: Located in the ruins northwest of The Mausoleum of King Deshret. The ruins can only be unlocked in The Secret of Al-Ahmar.
Dendroculus 139	Great Red Sand	Location: Located on a crumbled wall near an Exquisite Chest guarded by Primal Constructs.
Dendroculus 140	Great Red Sand	Location: Located north of The Mausoleum of King Deshret. Can only be unlocked in The Secret of Al-Ahmar World Quest.
Dendroculus 141	Great Red Sand	Location: Located near one of the Primal Torches during The Secret of Al-Ahmar World Quest. Complete the Primal Sandglass Puzzle in the room for a Four-Leaf Sigil to appear above, which will get you to the Dendroculus.
Dendroculus 142	Great Red Sand	Location: Gain enough clearance then fast travel to Khemenu Temple. Head west until you reach the main elevator room. In the room north, follow the path to find a plinth. Operate it to open the door and find the Dendroculus on the other side of the room.
Dendroculus 143	Great Red Sand	Location: Located in the buildings unlocked in An Introduction to Indoor Archaeology quest.
Dendroculus 144	Great Red Sand	Location: Located on the west end of Khemenu Temple's invisible ceiling.
Dendroculus 145	Great Red Sand	Location: Located outside of the The Mausoleum of King Deshret.
Dendroculus 146	Great Red Sand	Location: Located in front of a giant fan found while doing the Place of Breath objective of Dual Evidence World Quest.
Dendroculus 147	Great Red Sand	Location: Located in the buildings unlocked in An Introduction to Indoor Archaeology quest.
Dendroculus 148	Great Red Sand	Location: Located in the middle of a broken bridge in the Place of Breath Puzzle. Only accessible during the Dual Evidence World Quest and requires Lv. 2 clearance.
Dendroculus 149	Great Red Sand	Location: In the Place of Breath Puzzle ruins, accessed during Dual Evidence World Quest, move the primal beams to activate the two Anemo monuments to summon an upward draft at the center. The Dendroculus is at the top.
Dendroculus 150	Great Red Sand	Location: Located in The Mausoleum of King Deshret. Can only be unlocked after completing The Secret of Al-Ahmar World Quest.
Dendroculus 151	Great Red Sand	Location: Located in The Mausoleum of King Deshret. Can only be unlocked in The Secret of Al-Ahmar World Quest.
Dendroculus 152	Great Red Sand	Location: Located at the top of The Mausoleum of King Deshret.
Dendroculus 153	Great Red Sand	Location: Located in The Mausoleum of King Deshret. Can only be unlocked in The Secret of Al-Ahmar World Quest.
Dendroculus 154	Great Red Sand	Location: Found above the statues inside a cavern that can only be accessed by following The Secret of Al-Ahmar World Quest. Exit the room from the waypoint at the southwest corner of the mausoleum and use the Four-Leaf Sigil to get to the area above.
Dendroculus 155	Great Red Sand	Location: After obtaining Lv.3 clearance, solve the Primal Sandglass Puzzle on the other side and light up the Primal Torch with the ember to remove the invisible walls blocking this Dendroculus.
Dendroculus 156	Great Red Sand	Location: Activate the giant fan after gaining Lv.3 clearance and glide up to reach the Dendroculus.
Dendroculus 157	Great Red Sand	Location: Located on a tree in an underground Withering zone. the cavern can only be accessed by following The Secret of Al-Ahmar World Quest.
Dendroculus 158	Great Red Sand	Location: From the teleport waypoint by Dendroculus #50, face northwest and look for a tunnel above. Follow the path through it until you reach a room full of giant mushrooms. The Dendroculus is at the top of the biggest mushroom.
Dendroculus 159	Great Red Sand	Location: Find the entrance to the ruins on the southwest corner of the desert. Follow the path while solving the puzzles along the way and you will eventually run into this Dendroculus as you ascend through an elevator.
Dendroculus 160	Great Red Sand	Location: Located by the entrance to the ruins south of The Mausoleum of King Deshret. It is visible from the nearby teleport waypoint.
Dendroculus 161	Great Red Sand	Location: Gain enough clearance and enter The Dune of Magma. Find this Dendroculus in a pit beneath, just past a remarkable chest.
Dendroculus 162	Great Red Sand	Location: Located on a scaffolding that can be found while doing the Place of Breath objective of Dual Evidence World Quest. Interact with the rock pile at the base to summon an upward draft and reach the Dendroculus.
Dendroculus 163	Great Red Sand	Location: Located at the top of the pyramid in The Dune of Elusion. Use the Four-Leaf Sigil on the west and south walls to quickly reach the top.
Dendroculus 164	Great Red Sand	Location: Located inside the ruins south of The Dune of Elusion. The Dendroculus is found at the west end of the ruins, and will require at least Lv.3 clearance to complete.
Dendroculus 165	Great Red Sand	Location: Located south of The Dune of Elusion. Activate the three monuments with Pyro to lower the barrier surrounding the Dendroculus.
Dendroculus 166	Great Red Sand	Location: Located east of The Dune of Elusion, on a tree surrounded by Pyro Fungi.
Dendroculus 167	Great Red Sand	Location: Located south of Sobek Oasis. It is floating by a cliff east of a teleport waypoint.
Dendroculus 168	Great Red Sand	Location: Located southeast of Sobek Oasis. It is located above a Fatui tent.
Dendroculus 169	Great Red Sand	Location: Located north of the Valley of Dahri. It is above a pond in an underground cave north of a Statue of the Seven.
Dendroculus 170	Great Red Sand	Location: Located west of the Valley of Dahri. It is on a branch by tower.
Dendroculus 171	Great Red Sand	Location: Located inside an Eremite hideout. At the entrance, Paimon will start talking about tailing one of the eremites. Follow the Female Eremite walking down the road until she mentions a password. Use the password on the mechanism at the entrance to open the door. The Dendroculus is in the first room to the right.
Dendroculus 172	Great Red Sand	Location: Located by the Valley of Dahri. It is on a branch by a cliff. There is a trail of Four-Leaf Sigil by the teleport waypoint that can get you to the branches.
Dendroculus 173	Great Red Sand	Location: Located at the back of the Ruin Golem.
Dendroculus 174	Great Red Sand	Location: Located inside the Ruin Golem during the Afratu's Dilemma World Quest. The Dendroculus is on the wall next to a Suda's Flow Seal.
Dendroculus 175	Great Red Sand	Location: Located inside the Ruin Golem's right arm, behind a giant gear. The arm is accessible through the main floor.
Dendroculus 176	Great Red Sand	Location: After disabling the Ruin Golem's auto-fire during the Afratu's Dilemma World Quest. Take control of the machine and fire at one of the boulders on the right. Go through the opening to find a cavern with the Dendroculus inside.
Dendroculus 177	Great Red Sand	Location: Located in an underground cave with a Withering zone blocked by a boulder. The boulder can only be destroyed using the Ruin Golem by following the Afratu's Dilemma World Quest.
Dendroculus 178	Great Red Sand	Location: Located northeast of the Valley of Dahri. Use the Four-Leaf Sigil above the teleport waypoint to find and reach the Dendroculus.
Dendroculus 179	Great Red Sand	Location: Located in the ruins during the Lost in the Sands quest. Only accessible during the quest because the building entrance can only be unlocked from the inside.
Dendroculus 180	Great Red Sand	Location: Located above the temple entrance northeast of Sobek Oasis, next to the teleport waypoint.
Dendroculus 181	Desert of Hadramaveth	Location: Located behind a rock wall, near the Fatui Stronghold gate.
Dendroculus 182	Desert of Hadramaveth	Location: Located atop a ruined pillar, north of the waypoint.
Dendroculus 183	Desert of Hadramaveth	Location: Floating in the air, near the right arm of the Colossal Ruin Guard.
Dendroculus 184	Desert of Hadramaveth	Location: Located near an Eremite camp, under the Colossal Ruin Guard.
Dendroculus 185	Desert of Hadramaveth	Location: Located above the Debris of Panjvahe waypoint. Use the Four-Leaf Sigil to reach it.
Dendroculus 186	Desert of Hadramaveth	Location: Located near a ruin guard, north of the waypoint.
Dendroculus 187	Desert of Hadramaveth	Location: On top of a rock, near the fissure, in the Debris of Panjvahe.
Dendroculus 188	Desert of Hadramaveth	Location: Sealed behind a Dendro rock. Use Dendrograma to break them.
Dendroculus 189	Desert of Hadramaveth	Location: Located near an air draft, east of the waypoint.
Dendroculus 190	Desert of Hadramaveth	Location: Found floating across the Shrine of Depth.
Dendroculus 191	Desert of Hadramaveth	Location: Located by a high cliff just above the waypoint north of The Sands of Three Canals.
Dendroculus 192	Desert of Hadramaveth	Location: Located on top of a cactus northwest of The Sands of Three Canals.
Dendroculus 193	Desert of Hadramaveth	Location: Located in the same area as the Primal Construct battle arena of the Dune-Emtombed Fecundity: Part 2.
Dendroculus 194	Desert of Hadramaveth	Location: Located high above the air at The Sands of Three Canals. Activate the Anemo Monument to summon an updraft. Glide up until you spot the Four-Leaf Sigil east of your location. Launch yourself then glide towards the Dendroculus until you land on an invisible platform. From there, keep running up then jump to acquire it.
Dendroculus 195	Desert of Hadramaveth	Location: Sealed behind a Dendro rock. Use Dendrograna to break them.
Dendroculus 196	Desert of Hadramaveth	Location: Located right below the waypoint west of The Sands of Three Canals hanging by a branch.
Dendroculus 197	Desert of Hadramaveth	Location: Located on top of a dead tree at The Sands of Three Canals.
Dendroculus 198	Desert of Hadramaveth	Location: Located inside a tent south of The Sands of Three Canals.
Dendroculus 199	Desert of Hadramaveth	Location: Sealed inside the Dendro rock. Head to the Dendrograma branch surrounded by an Atmospheric Vortex and Geo slimes. Equip your Liloupar gadget to unlock it.
Dendroculus 200	Desert of Hadramaveth	Location: Located floating in the air high above the valley west of The Sands of Three Canals. Position yourself on the cliff by the huge branch, parallel to the path of the Dendroculus. Run up and an invisible bridge will appear that you can use to get close. Jump to acquire it.
Dendroculus 201	Desert of Hadramaveth	Location: Located on top of a dead tree at Wounded Shin Valley.
Dendroculus 202	Desert of Hadramaveth	Location: Located inside the hollow space of the head of the Ruin Golem. This is locked behind the Dune-Entombed Fecundity: Part 3 quest of The Dirge of Bilqis.
Dendroculus 203	Desert of Hadramaveth	Location: Found in the area where you obtain Liloupar's fragment during the Dune-Entombed Fecundity: Part 3 quest of The Dirge of Bilqis.
Dendroculus 204	Desert of Hadramaveth	Location: Located deep inside a small cave in the Ruin Golem tunnels. This is locked behind the Dune-Entombed Fecundity: Part 3 quest of The Dirge of Bilqis.
Dendroculus 205	Desert of Hadramaveth	Location: Located on top of a dead tree between Wounded Shin Valley and The Temples Forsaken.
Dendroculus 206	Desert of Hadramaveth	Location: Located inside the small temple northwest of The Temples Forsaken, floating by the Four-Leaf Sigil
Dendroculus 207	Desert of Hadramaveth	Location: Located next to the waypoint west of Safhe Shatranj by the Four-Leaf Sigil.
Dendroculus 208	Desert of Hadramaveth	Location: Obtainable after the sand recedes for completing the ruins in The Temple Where Sand Flows Like Tears quest of The Dirge of Bilqis.
Dendroculus 209	Desert of Hadramaveth	Location: Located outside The Temples Forsaken, near the east waypoint.
Dendroculus 210	Desert of Hadramaveth	Location: Located inside a small cave outside The Temple Forsaken.
Dendroculus 211	Desert of Hadramaveth	Location: Located inside the small cave right above the Four-Leaf Sigil.
Dendroculus 212	Desert of Hadramaveth	Location: Located on top of a dead tree at the Tanit Camps.
Dendroculus 213	Desert of Hadramaveth	Location: Located on top of a broken pillar beneath the waypoint east of Wadi Al-Majuj.
Dendroculus 214	Desert of Hadramaveth	Location: In the underground area northwest of Wadi Al-Majuj, near the top of a large statue.
Dendroculus 215	Desert of Hadramaveth	Location: Located inside the ruins after completing the Apocalypse Lost quest. You must go up an invisible ramp.
Dendroculus 216	Desert of Hadramaveth	Location: Located hanging on a branch north of Wadi Al-Majuj.
Dendroculus 217	Desert of Hadramaveth	Location: Located on top of a broken pillar beneath the waypoint east of Wadi Al-Majuj.
Dendroculus 218	Desert of Hadramaveth	Location: Located near the center of the Eternal Oasis. This is locked behind The Eternal Dream, Ever Lush quest of The Dirge of Bilqis.
Dendroculus 219	Desert of Hadramaveth	Location: Located by the grassy area of the river shore. This is locked behind The Eternal Dream, Ever Lush quest of The Dirge of Bilqis.
Dendroculus 220	Desert of Hadramaveth	Location: Located inside an Atmospheric Vortex. Equip your max level Liloupar gadget upgraded from Tadhla's quest chain to unlock it.
Dendroculus 221	Desert of Hadramaveth	Location: Located atop a tree, above an Eremite camp, in Passage of Ghouls.
Dendroculus 222	Desert of Hadramaveth	Location: Floating on a large rock, down the fissure in Passage of Ghouls.
Dendroculus 223	Desert of Hadramaveth	Location: Located on top of the tree, south of the Qusayr Al-Inkhida' waypoint.
Dendroculus 224	Desert of Hadramaveth	Location: Use the Four-Leaf Sigil to get to the top of the room. The waypoint is locked behind the Dune-Entombed Fecundity: Part 1 quest of The Dirge of Bilqis.
Dendroculus 225	Desert of Hadramaveth	Location: Located high in the air. Use Dendrograma to break the Dendro rocks and reveal the Dendro monuments. Light all four monuments to activate a wind current. Glide up to get the Dendroculus.
Dendroculus 226	Desert of Hadramaveth	Location: Found hidden inside the tree trunk inside the Eternal Oasis. This is locked behind The Eternal Dream, Ever Lush quest of The Dirge of Bilqis.
Dendroculus 227	Desert of Hadramaveth	Location: Found on top of a cactus, south of The Sands of Three Canals.
Dendroculus 228	Desert of Hadramaveth	Location: Sealed behind a Dendro rock. Use Dendrograma to break them.
Dendroculus 229	Desert of Hadramaveth	Location: Located floating above the Setekh Wenut tunnel entrance.
Dendroculus 230	Desert of Hadramaveth	Location: Underground, near the City of Deceased domain, hidden behind a pile of rocks.
Dendroculus 231	Desert of Hadramaveth	Location: Investigate a rock formation located inside the Wenut Tunnels to reveal a hole. Go to the northern ledge and look into the hole.
Dendroculus 232	Desert of Hadramaveth	Location: Follow the The Temple Where Sand Flows Like Tears quest of The Dirge of Bilqis and get to the second Ruin Mechanism room. It is found atop of a large branch.
Dendroculus 233	Desert of Hadramaveth	Location: Blow off the sand pile on the mechanism and activate it to reveal the hole. This is located during The Temple Where Sand Flows Like Tears quest of The Dirge of Bilqis
Dendroculus 234	Desert of Hadramaveth	Location: Begin The Temple Where Sand Flows Like Tears quest of The Dirge of Bilqis to get inside the temple. Follow the quest and solve the Rune Mechanism puzzles to see the Dendroculus.
Dendroculus 235	Desert of Hadramaveth	Location: Located inside the cavern, atop the branches inside a ruin door, during the Dune-Entombed Fecundy: Part 2.
Dendroculus 236	Girdle of the Sands	Location: On top of a hut in a Hilichurl camp west of the waypoint.
Dendroculus 237	Girdle of the Sands	Location: On a cactus above the Molten Iron Fortress Domain.
Dendroculus 238	Girdle of the Sands	Location: At the tip of a pole next to an Anemo Hilichurl Rogue.
Dendroculus 239	Girdle of the Sands	Location: In an underground area at Temir Mountains. The Dendroculus is found near the bottom of the shaft.
Dendroculus 240	Girdle of the Sands	Location: At the tip of a gray crystal.
Dendroculus 241	Girdle of the Sands	Location: On top of a large forge found during the Khvarena of Good and Evil quest.
Dendroculus 242	Girdle of the Sands	Location: Along the edge of the cliff. Use the Four-Leaf Sigils to quickly gain elevation then climb the cliff to reach the Dendroculus.
Dendroculus 243	Girdle of the Sands	Location: At the tip of one of the large gray crystals surrounding Tunigi Hollow.
Dendroculus 244	Girdle of the Sands	Location: Underground by a cliff with some Four-Leaf Sigils nearby. Requires progression through the Khvarena of Good and Evil quest.
Dendroculus 245	Girdle of the Sands	Location: At the center of Tunigi Hollow.
Dendroculus 246	Girdle of the Sands	Location: Under the elevator after connecting the three Mysterious Energy Receiver during the Khvarena of Good and Evil quest.
Dendroculus 247	Girdle of the Sands	Location: On a large gray crystal east of Tunigi Hollow. It's hard to reach using the nearby sigil, so you'll have to get on top of the east cliff then glide down.
Dendroculus 248	Girdle of the Sands	Location: Near the ceiling inside a room with wind currents and sunlight. Requires progression through the Khvarena of Good and Evil quest.
Dendroculus 249	Girdle of the Sands	Location: Under an arch just below a Four-Leaf Sigil.
Dendroculus 250	Girdle of the Sands	Location: Above a tree that's high up a cliff with a Consecrated Red Vulture nearby.
Dendroculus 251	Girdle of the Sands	Location: Found inside a tree stump.
Dendroculus 252	Girdle of the Sands	Location: Floating in the air near a high cliff. Requires you to use Sorush to adjust the Four-Leaf Sigil.
Dendroculus 253	Girdle of the Sands	Location: On top of one of the giant tree roots.
Dendroculus 254	Girdle of the Sands	Location: Floating in the air between some branches. Climb to the highest point you can get on the tree and glide your way to the Dendroculus.
Dendroculus 255	Girdle of the Sands	Location: Floating in the air. Climb the nearby cliff and glide down to it.
Dendroculus 256	Girdle of the Sands	Location: Floating on top of a rock. Get onto a higher ground then glide down to reach it.
Dendroculus 257	Girdle of the Sands	Location: Floating beneath a small rock bridge, in an opening that leads undeground.
Dendroculus 258	Girdle of the Sands	Location: On top of a small tree next to a Soul Bell.
Dendroculus 259	Girdle of the Sands	Location: Sitting on top of a Treasure Hoarder hut by the cliff.
Dendroculus 260	Girdle of the Sands	Location: On top of a large tree.
Dendroculus 261	Girdle of the Sands	Location: On top of a large branch.
Dendroculus 262	Girdle of the Sands	Location: Found above the water. Glide from one of the cliffs in the Samudra Coast.
Dendroculus 263	Girdle of the Sands	Location: Inside an underground arena found while progressing through the Khvarena of Good and Evil quest.
Dendroculus 264	Girdle of the Sands	Location: Underground in an area full of Mysterious Energy Control Systems. Take the elevator up to an area with two Ruin Guards to find the Dendroculus.
Dendroculus 265	Girdle of the Sands	Location: Found behind a gray crystal entrance inside an underground cave.
Dendroculus 266	Girdle of the Sands	Location: Found between the top of a large gray crystal and a cliff.
Dendroculus 267	Girdle of the Sands	Location: Beneath the edge of a large cliff.
Dendroculus 268	Girdle of the Sands	Location: On top of a large spike in the underground cave at the Gate of Zulqarnain, following the road headed north.
Dendroculus 269	Girdle of the Sands	Location: Found beneath a large cliff with a Consecrated Red Vulture on top.
Dendroculus 270	Girdle of the Sands	Location: Found on top of a hut in a Hilichurl camp west of the Gate of Zulqarnain waypoint.
Dendroculus 271	Girdle of the Sands	Location: Found inside a broken piece of machinery next to some Ruin Drakes.

There are a total of 271 Dendroculi that you can collect in the region of Sumeru. 224 of them are discovered on the surface, and 47 are found underground!

Sumeru Map and Unlock Guide

All Dendroculi Locations

Rainforest Dendroculus Open Image	Great Red Sand Dendroculus Open Image
Hadramaveth Dendroculus Open Image	Girdle of the Sands Open Image

How to Use the Dendroculus

Upgrade the Sumeru Statue of the Seven

Genshin - Sumeru Teaser Statue of the Seven

Dendroculus is a collectible item that can be found in the Sumeru region. You use them to level up the Statues of the Seven in Sumeru, which rewards you with Primogems, Shrine of Depths Keys, and Adventure EXP.

Extra Dendroculus

As per usual, 1 Dendroculus is expected to be left in your inventory. There are no known uses yet for the extra Oculi.

How to Level Up Statues of the Seven

How Many Dendroculus are There?

There are 271 Dendroculus total found in the Sumeru region!

Search with Friends

If you're finding it hard to explore the terrains of Sumeru, bring a friend or two to help you! Check out our friend request and co-op boards for potential exploration buddies:

Genshin Co-Op Boards
Friend Request Board	Co-Op Board	Supporting Cast Board

Rewards from the Sumeru Statue of the Seven

Lv.	Dendroculus Required	Rewards
2	20	Adventure EXP x80 Primogem x60 Dendro Sigil x5 Sumeru Shrine of Depths Key x1
3	22	Adventure EXP x120 Primogem x60 Dendro Sigil x5 Memory of Flourishing Green x1
4	25	Adventure EXP x160 Primogem x60 Dendro Sigil x5 Sumeru Shrine of Depths Key x1
5	28	Adventure EXP x200 Primogem x60 Dendro Sigil x10 Memory of Flourishing Green x1
6	30	Adventure EXP x240 Primogem x60 Dendro Sigil x10 Sumeru Shrine of Depths Key x1
7	32	Adventure EXP x280 Primogem x60 Dendro Sigil x10 Memory of Flourishing Green x1
8	35	Adventure EXP x320 Primogem x60 Dendro Sigil x15 Sumeru Shrine of Depths Key x1
9	38	Adventure EXP x360 Primogem x60 Dendro Sigil x15 Sumeru Shrine of Depths Key x1
10	40	Adventure EXP x400 Primogem x60 Dendro Sigil x15 Sumeru Shrine of Depths Key x1

Note that players who have not reached Adventure Rank 60 can still gain Adventure EXP instead of Mora from the Statue of the Seven.

Genshin Impact Related Guides

Genshin Impact - Areas and Map Guides

All Interactive Map Guides

Map Guides for Teyvat Nations

List of Region Map Guides
Mondstadt Map Guide	Liyue Map Guide
Inazuma Map Guide	Sumeru Map Guide
Fontaine Map Guide	Natlan Map Guide
Nod-Krai Map Guide

All Nations and Regions

Map Guides for Other Areas

List of Map Guides
Enkanomiya	Golden Archipelago
Dragonspine	The Chasm
Sumeru Rainforest	Sumeru Desert
Veluriyam Mirage	Chenyu Vale
Simulanka	Sacred City of Tollan
Easybreeze Holiday Resort

Oculi Location Guides

Teyvat Oculus Guides
All Oculi	Anemoculus	Geoculus	Electroculus
Dendroculus	Hydroculus	Pyroculus	Lunoculus

Other Oculus Map Guides

Dendroculus Guides
Rainforest Dendroculus	All Desert Dendroculus
Hadramaveth Dendroculus	Girdle of the Sands Dendroculus

Comment

Jesse4games◆AhcOQ..whI 4 months

Thanks for your help with everything especially at this time with Dendroculus. Challenge completed.

Anonymous over 2 years

There appear to be three that are locked in quests. I've already completed them, but I think I didn't collect the dendruculus-s during them, didn't notice they were there or thought I could go back. Is there no way to make up for that? If there's only one leftover, I think that means I can't ever fully upgrade the statue.

See more comments (22 Comments)

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Dendroculus Locations and Interactive Map

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Dendroculus Locations Map

All Dendroculi Locations

How to Use the Dendroculus

Upgrade the Sumeru Statue of the Seven

Extra Dendroculus

How Many Dendroculus are There?

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